VERONA PHARMA PLC: Holding(s) in Company

 For filings with the FCA include the annex                                                                                                                                           For filings with issuer exclude the annex                                                                                                                                            TR-1: NOTIFICATION OF MAJOR INTEREST IN SHARESi                                                        1. Identity of the issuer or the underlying issuer                                     of existing shares to which voting rights are                        Verona  Pharma plc   attached:ii                                                                                                                                                                          2. Reason for the notification (please tick the appropriate box or boxes):                                                                                                            An acquisition or disposal of voting rights                                                √                                                                                         An acquisition or disposal of qualifying financial instruments which may result        in the acquisition of shares already issued to which voting rights are attached                                                                                                      An acquisition or disposal of instruments with similar economic effect to              qualifying financial instruments                                                                                                                                                     An event changing the breakdown of voting rights                                                                                                                                     Other (please specify):  CHANGE TO THE NUMBER OF VOTING RIGHTS IN ISSUE,  FOLLOWING THE √                             RECENT SUBSCRIPTION, PLACING AND OPEN OFFER.                                                                                                             3. Full name of person(s)                                                              subject to the            Aviva plc & its subsidiaries                                 notification obligation:                                                               iii                                                                                                                                                                                                              Registered Holder:                                                                                                                                                                                                                                                            BNY Norwich Union Nominees Limited     4,817,119*                                                                                                                                                                                                                              Chase (GA Group) Nominees Limited    139,178,649*             4. Full name of                                                                        shareholder(s)            *denotes direct interest                                      (if different from 3.):                                                               iv                                                                                                                     Chase (GA Group) Nominees Limited      15,111,869                                                                                                                                                                                                                             Chase Nominees Limited                  4,440,952                                                                                                                                                                                                               Vidacos Nominees Limited               18,701,411                                                                                                                                                                                                                                                                               5. Date of the                                                                         transaction and date on                                                                which the threshold is    25 March 2014                                                crossed or                                                                             reached: v                                                                                                                                                                           6. Date on which issuer   27 March 2014                                                notified:                                                                                                                                                                            7. Threshold(s) that is/  0% to 14% Change at Direct Interest Level and                are crossed or                                                                         reached: vi, vii          0% to 18% Change at Combined Interest Level                                                                                                                    8. Notified details:                                                                                                                                                              A: Voting rights attached to shares viii, ix                                                                                                                                         Class/type of   Situation         Resulting situation after the triggering  transaction   shares          previous                                                                                     to the triggering                                                                         transaction                                                             if possible                                                                            using           Number   Number   Number      Number of voting       % of   voting rights the ISIN CODE   of       of       of shares   rights                 x                                       Shares   Voting                                                                                  Rights   Direct      Direct xi   Indirect   Direct    Indirect                                                                xii                                                                                                                     Ordinary Shares                                                                                                                                                                                      NIL HELD NIL HELD 182,250,000 143,995,768 38,254,232 14.26%    3.79%      GB00B06GSH43                                                                                                                                                                                                                                                                       B: Qualifying Financial Instruments                                                                                                                                               Resulting situation after the triggering transaction                                                                                                                                 Type of financial  Expiration Exercise/              Number of voting       %  of voting  instrument         date xiii  Conversion Period xiv  rights that may be      rights                                                               acquired if the                                                                           instrument is                                                                             exercised/ converted.                                                                                                                                                                                                      C: Financial Instruments with similar economic effect to Qualifying Financial           Instruments xv, xvi                                                                                                                                                                  Resulting situation after the triggering transaction                                                                                                                                 Type of    Exercise Expiration   Exercise/     Number of voting rights   % of  voting     financial  price    date xvii    Conversion    instrument refers to      rights  xix, xx  instrument                       period xviii                                                                                                                                                                                                                                                                                                            Nominal   Delta                                                                                                                                                                                          Total (A+B+C)                                                                                                                                                                     Number of voting rights                  Percentage of voting rights                                                                                                                 182,250,000                              18.05%                                                                                                                                          9. Chain of controlled undertakings through which the voting rights and/or the          financial instruments are effectively held, if applicable: xxi                                                                                                                       The voting rights are managed and controlled by Aviva Investors Global Services        Limited, with the following chain of controlled undertakings:-                                                                                                                       Aviva Investors Global Services Limited:                                                                                                                                             Aviva plc (Parent Company)                                                                                                                                                           Aviva Group Holdings Limited (wholly owned subsidiary of Aviva plc)                                                                                                                  Aviva Investors Holdings Limited (wholly owned subsidiary of Aviva Group               Holdings Limited)                                                                                                                                                                    Aviva Investors Global Services Limited (wholly owned subsidiary of Aviva              Investors Holdings Limited)                                                                                                                                                                                                                                                             Proxy Voting:                                                                                                                                                                     10. Name of the proxy holder:                                    See Section 4                                                                                           11. Number of voting rights proxy holder will cease                                    to hold:                                                                                                                                                                             12. Date on which proxy holder will cease to hold                                      voting rights:                                                                                                                                                                                                                                                                 13. Additional   Figures are based on a total number of voting rights of  1,009,923,481,  information:     as per the Company's Result of General Meeting Announcement of  24 March                     2014.                                                                                                                                                                                                                                                      14. Contact      Neil Whittaker, Aviva plc                                              name:                                                                                                                                                                             15. Contact                                                                            telephone        01603 684420                                                           number:                                                                                                                                                                                 END  -0- Mar/27/2014 12:25 GMT    
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